Department Chair
Scott M. Boback
Associate Professor of Biology (2007).
Rector North Room 1303
(717) 245-1799
Department Faculty
John H. Henson
Senior Associate Provost for Academic Affairs; Charles A. Dana Professor of Biology (1989).
West College (Old West) Room 8
(717) 245-1434 |
B.A., University of Virginia, 1979; M.S., Florida State University, 1983; Ph.D., Harvard University, 1989.

He teaches courses in cell biology, immunology, animal development, marine science, and health studies. His research program utilizes marine orgnisms as model experimental systems for studying basic aspects of cell structure and function, particularly how cells move, change shape, and divide.
Thomas M. Arnold
Professor of Biology (2003).
Rector North Room 2303
(717) 245-1319 |
B.A., St. Mary's College of Maryland, 1993; Ph.D., University of Delaware, 1998.

Dr. Arnold is a biochemist and physiologist who studies natural toxins, pheromones, odors, and anti-microbials. He focuses on natural products found in both marine and terrestrial ecosystems, including seagrass communities, coral reefs, temperate forests, and agricultural fields.
Anthony Pires
Professor of Biology (1993).
James Hall - Rector Complex Room 1225
(717) 245-1632 |
B.A., Harvard College, 1982; Ph.D., Cornell University, 1990.

He teaches courses in neurobiology, behavioral biology and invertebrate zoology. His research interests are in the neural control of developmental events, especially the regulation of settlement and metamorphosis in larvae of marine invertebrates.
Carol C. Loeffler
Associate Professor of Biology (1988).
Rector North Room 2304
(717) 245-1360 |
B.A., Smith College, 1982; Ph.D., Cornell University, 1992.
Dickinson Award for Distinguished Teaching, 2008-09.

She teaches courses in algae, fungi, lichens, and land plants. Her research interests with students are in the biology and ecology of rare plant species and in the impact of deer and other herbivores on forest vegetation.
Michael P. Roberts
Associate Professor of Biology (1992).
Rector North Room 1304
(717) 245-1201 |
B.A., Colgate University, 1977; M.S., Miami University, 1979; Ph.D., Yale University, 1988.
Ganoe Award for Inspirational Teaching, 2002-03.

His research interests center on the biology of cancer, in particular leukemia, and the mechanisms by which cancer cells might be reprogrammed to either behave normally or self destruct. He and his students use genomic, proteomic and bioinformatic methods to explore changes in gene expression that distinguish cancer cells from normal cells. His teaching interests included genetics, molecular genetics, bioinformatics and the biology of cancer.
Charles F. Zwemer
Associate Professor of Biology (1995).
James Hall - Rector Complex Room 1217
(717) 245-1293 |
B.A., Hope College, 1987; Ph.D., Indiana University Medical Sciences Program, 1993; Post-Doctoral Fellowship, University of Michigan Medical School, 1995.
Ganoe Award for Inspirational Teaching, 1999-2000.

He teaches courses in physiology, microanatomy, and vertebrate biology. His research addresses issues of cardiovascular, respiratory, and renal function in normal and diseased states.
Kirsten A. Guss
Associate Professor of Biology; John R. & Inge Paul Stafford Chair in Bioinformatics (2001).
Rector North Room 2302
(717) 245-1530 |
B.A., Gettysburg College, 1989; Ph.D., Carnegie Mellon University, 1997.

Prof. Guss is interested in the molecular basis of development. Her research focuses on the role of the transcription factor scalloped during development of the fruit fly Drosophila melanogaster. Prof. Guss teaches the following courses in the biology program: 127: This is Your Life, 317: Genetics, 318: Animal Development, and 418: Developmental Genomics.
David B. Kushner
(Director, Norwich Science Program in England, 2015-17)
Associate Professor of Biology (2003).
Rector North Room 1302
(717) 245-1328 |
B.S., Haverford College, 1993; Ph.D., University of Pennsylvania, 1998.

He teaches courses in microbiology, virology, and RNA biology. His research combines genetics and cell and molecular and biology with modern genomic and bioinformatic approaches to understand interactions between viruses and their hosts, and also uses evolution-based approaches to understand the relationship between viral RNA sequence, structure, and function.
Scott M. Boback
Associate Professor of Biology (2007).
Rector North Room 1303
(717) 245-1799 |
B.A., Ripon College, 1991; M.A., University of Northern Colorado, 1995; Ph.D., Auburn University, 2005.

Broadly, I am an animal ecologist with a focus in herpetology (non-avian reptiles and amphibians). I teach courses in ecology, evolution, vertebrate natural history, and physiology. My students and I investigate a variety of organisms in both lab and field settings. Recent projects have included the muscular performance of snakes during constriction, the population ecology of painted turtles, and the evolutionary development of snake jaws.
Mary M. Niblock
Associate Professor of Biology (2007).
James Hall - Rector Complex Room 1221
(717) 245-1292 |
B.A., University of Richmond, 1992; Ph.D., Wake Forest University, 1998.

Tiffany A. Frey
Assistant Professor of Biology (2011).
James Hall - Rector Complex Room 2221
(717) 254-8037 |
B.S., The Pennsylvania State University, 2000; Ph.D., Johns Hopkins University School of Medicine, 2008.

Dana J. Wohlbach
Assistant Professor of Biology (2013).
James Hall - Rector Complex Room 2219
(717) 254-8131 |
B.A., Franklin and Marshall College, 2002; Ph.D., University of Wisconsin-Madison, 2007.

Jennifer Wanat
Visiting Assistant Professor of Biology (2015).
Rector North Room 1302
(717) 254-8333 |
B.S., University of Rhode Island, 2001; Ph.D., Cornell University, 2007.

H. Eugene Wingert
Visiting Instructor in Biology (2007).
Rector North Room 1310
(717) 254-8939 |
B.A., Shippensburg University, 1967; M.Ed., 1972.

Adjunct Faculty
Meredith Rauhut
(Leave of absence 2016-17)
Instructor in Neuroscience (Spring 2015).
James Hall - Rector Complex Room 1219
(717) 254-8193 |
B.A., Bucknell University, 1995; Ph.D., University of Massachusetts-Amherst, 2001.

Jason Smith
Adjunct Faculty in Biology (2016).
James Hall - Rector Complex Room 1223
(717) 254-8344 |

Academic Administrative Staff
Michael Potthoff
Department Technician
Rector North Room 1308
(717) 245-1144 |

Emeriti Faculty
Thomas Brennan
Joseph Priestley Professor Emeritus
417 South College Street
Carlisle, PA 17013
717-243-5015 (Home) |

Janet Wright